The pair will join Macbeth duo Michael Fassbender and Marion Cotillard in the upcoming video game adaptation, Deadline reports.
Irons will play Cotillard’s father, while Gleeson will take on the role of Fassbender’s father in the film.
The film follows the same plot premise as the games: by unlocking his genetic code, Fassbender’s protagonist Callum Lynch is able to relive his ancestor’s memories of jumping over buildings and stabbing people in days of yore.
The film, directed by Macbeth helmer Justin Kurzel, began production in August.
The hugely popular Assassin’s Creed gaming series continues with Assassin’s Creed Syndicate, which launches on October 23 for PlayStation 4 and Xbox One.
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